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WhatDoWhalesFeel?Anexaminationofthefunctioningofthesensesincetaceans,thegroupofmammalscomprisingwhales,dolphinsandporpoisesSomeofthesensesthatweandotherterrestrialmammalstakeforgrantedareeitherreducedorabsentincetaceansorfailtofunctionwellinwater.Forexample,itappearsfromtheirbrainstructurethattoothedspeciesareunabletosmell.Baleenspecies,ontheotherhand,appeartohavesomerelatedbrainstructuresbutitisnotknownwhetherthesearefunctional.Ithasbeenspeculatedthat,astheblowholesevolvedandmigratedtothetopofthehead,theneuralpathwaysservingsenseofsmellmayhavebeennearlyallsacrificed.Similarly,althoughatleastsomecetaceanshavetastebuds,thenervesservingthesehavedegeneratedorarerudimentary.Thesenseoftouchhassometimesbeendescribedasweaktoo,butthisviewisprobablymistaken.Trainersofcaptivedolphinsandsmallwhalesoftenremarkontheiranimals’responsivenesstobeingtouchedorrubbed,andbothcaptiveandfreerangingcetaceanindividualsofallspecies(particularlyadultsandcalves,ormembersofthesamesubgroup)appeartomakefrequentcontact.Thiscontactmayhelptomaintainorderwithinagroup,andstrokingortouchingarepartofthecourtshipritualinmostspecies.Theareaaroundtheblowholeisalsoparticularlysensitiveandcaptiveanimalsoftenobjectstronglytobeingtouchedthere.Thesenseofvisionisdevelopedtodifferentdegreesindifferentspecies.Baleenspeciesstudiedatclosequartersunderwater–specificallyagreywhalecalfincaptivityforayear,andfree-rangingrightwhalesandhumpbackwhalesstudiedandfilmedoffArgentinaandHawaii–haveobviouslytrackedobjectswithvisionunderwater,andtheycanapparentlyseemoderatelywellbothinwaterandinair.However,thepositionoftheeyessorestrictsthefieldofvisioninbaleenwhalesthattheyprobablydonothavestereoscopicvision.Ontheotherhand,thepositionoftheeyesinmostdolphinsandporpoisessuggeststhattheyhavestereoscopicvisionforwardanddownward.Eyepositioninfreshwaterdolphins,whichoftenswimontheirsideorupsidedownwhilefeeding,suggeststhatwhatvisiontheyhaveisstereoscopicforwardandupward.Bycomparison,thebottlenosedolphinhasextremelykeenvisioninwater.Judgingfromthewayitwatchesandtracksairborneflyingfish,itcanapparentlyseefairlywellthroughtheair–waterinterfaceaswell.Andalthoughpreliminaryexperimentalevidencesuggeststhattheirin-airvisionispoor,theaccuracywithwhichdolphinsleaphightotakesmallfishoutofatrainer’shandprovidesanecdotalevidencetothecontrary.Suchvariationcannodoubtbeexplainedwithreferencetothehabitatsinwhichindividualspecieshavedeveloped.Forexample,visionisobviouslymoreusefultospeciesinhabitingclearopenwatersthantothoselivinginturbidriversandfloodedplains.TheSouthAmericanboutuandChinesebeiji,forinstance,appeartohaveverylimitedvision,andtheIndiansususareblind,theireyesreducedtoslitsthatprobablyallowthemtosenseonlythedirectionandintensityoflight.Althoughthesensesoftasteandsmellappeartohavedeteriorated,andvisioninwaterappearstobeuncertain,suchweaknessesaremorethancompensatedforbycetaceans’well-developedacousticsense.Mostspeciesarehighlyvocal,althoughtheyvaryintherangeofsoundstheyproduce,andmanyforageforfoodusingecholocation.Largebaleenwhalesprimarilyusethelowerfrequenciesandareoftenlimitedintheirrepertoire.Notableexceptionsarethenearlysong-likechorusesofbowheadwhalesinsummerandthecomplex,hauntingutterancesofthehumpbackwhales.Toothedspeciesingeneralemploymoreofthefrequencyspectrum,andproduceawidervarietyofsounds,thanbaleenspecies(thoughthespermwhaleapparentlyproducesamonotonousseriesofhigh-energyclicksandlittleelse).Someofthemorecomplicatedsoundsareclearlycommunicative,althoughwhatroletheymayplayinthesociallifeand‘culture’ofcetaceanshasbeenmorethesubjectofwildspeculationthanofsolidscience.Questions15-21Completethetablebelow.ChooseNOMORETHANTHREEWORDSfromReadingPassage2foreachanswer.Writeyouranswersinboxes15–21onyouranswersheet.SENSESPECIESABILITYCOMMENTSSmelltoothednoevidencefrombrainstructurebaleennotcertainrelatedbrainstructuresarepresentTastesometypespoornerveslinkedtotheir15....................areunderdevelopedTouchallyesregionaroundtheblowholeverysensitiveVision16....................yesprobablydonothavestereoscopicvisiondolphins,porpoisesyesprobablyhavestereoscopicvision17....................and....................18....................yesprobablyhavestereoscopicvisionforwardandupwardbottlenosedolphinyesexceptionalin19....................andgoodinair-waterinterfaceboutuandbeijipoorhavelimitedvisionIndiansusunoprobablyonlysensedirectionandintensityoflightHearingmostlargebaleenyesusuallyuse20....................repertoirelimited21....................whalesand....................whalesyessong-liketoothedyesusemoreoffrequencyspectrum;havewiderrepertoireQuestions22-26AnswerthequestionsbelowusingNOMORETHANTHREEWORDSfromthepassageforeachanswer.Writeyouranswersinboxes22–26onyouranswersheet.22Whichofthesensesisdescribedhereasbeinginvolvedinmating?23Whichspeciesswimsupsidedownwhileeating?24Whatcanbottlenosedolphinsfollowfromunderthewater?25Whichtypeofhabitatisrelatedtogoodvisualability?26Whichofthesensesisbestdevelopedincetaceans?
本文标题:C4T1P2-how-do-whales-feel
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